Frog

Frog is the common name for amphibians in the order, Anura. There is often a distinction made between frogs and toads, based on appearance, however there is no scientific distinction. The only family exclusively given the common name "toad" is Bufonidae, but many species from various other families are also called "toads". more...

"True frogs" are of the family Ranidae.

Characteristics

Due to the large diversity of frogs (over 4000 species), many characteristics are not shared throughout all of the species. However, there are some characteristics which distinguish them from other amphibians. Frogs are generally well suited to jumping, and have long hind legs, with elongated ankle bones. They have a short vertebral column, with no more than ten free vertebrae, followed by a fused tail bone, usually resulting in a frog with no tail.

Frogs range in size from 10mm (Psyllophryne didactyla of Brazil and Eleutherodactylus iberia of Cuba) to 300mm (Goliath frog, Conraua goliath, of Cameroon).

The skin is unkeratinized and hangs loosely on the body because of the lack of loose connective tissue, and can be smooth, warty or have skin folds. Frogs have three eyelid membranes: one transparent to protect the eyes underwater, and two which are translucent to opaque. They have a tympanum on each side of their head, which is involved in hearing, and is covered by skin in some species.

Call

The males of most species of frogs will call for a variety of reasons. It will call by passing air through the larynx, in the throat. In most calling frogs, the sound is amplified by the vocal sac(s). The vocal sac is a membrane of skin under the throat which distends during the amplification of the call. Each call is unique to a species.

The main reason for calling is for the male to attract a mate. Males will either call in a group, called a chorus, or individually. A male frog will emit a different call when mounted by another male. Both of these calls are emitted with the mouth of the frog closed.

A distress call is emitted by some frogs when they are in a position of danger. This is done with the mouth open, and usually results in a higher pitched call. The effectiveness of the call is unknown, however it is suspected the call intrigues the predator, until another animal is attracted, distracting them enough for its escape.

Many species of frog have deep calls, or croaks. Frog noise tends to be spelt (for English speakers) as "crrrrk" in Britain and "ribbit" in the USA. This difference is due to the different species within each region(e.g. Common frog (Rana temporaria) in Britain and Leopard frog (Rana pipiens) in the USA). The croak of the American bullfrog (Rana catesbiana) is sometimes spelt "jug o' rum".

Poison

Some species of frog secrete toxins from their skin. These toxins deter predatory animals from eating them, and some are extremely poisonous to humans. Generally the frog obtains the poison from the insects or animals they eat. The Australian corroboree frogs pseudophryne corroboree and pseudophryne pengilleyi have been discovered to be able to manufacture a unique alkaloid which is not derived from their diet (Daly). Some natives of the Amazon area extract poison from the poison dart frog and put it on their darts when hunting. It was previously a misconception that the poison was placed on arrows rather than darts. The name of the frog was changed from poison arrow frog to poison dart frog in the early 1980's. Poisonous frogs tend to advertise their toxicity with bright colours.

Distribution and status

Frogs are found nearly worldwide, but they do not occur in Antarctica and are not present on many oceanic islands.

In many parts of the world, the frog populations have declined drastically since the 1950s. Many environmental scientists feel that amphibians, and frogs in particular, may be excellent biological indicators of ecosystem function because of their location on the food web, their permeable skins and their typically bi-phasic life (in both water and on land). Although habitat loss is certainly one of the most important features of most declines; pollutants, climate change, introduction of non-indigenous predators/competitors, and infectious diseases (see Chytrid fungus) have also been implicated.

Life cycle

The life cycle of frogs contains many stages. Typically, the male frogs of a species will assemble at a still water source. They will then call, collectively becoming a chorus of frogs. The call is unique to the species, and will attract females of that species.

The male and female frog, will then undergo amplexus. This involves the male mounting the female. The female then releases her eggs, which the male frog covers with a sperm solution before the eggs make contact with the water. Once the eggs come in contact with the water, they will swell, and form a protective coating. The eggs are typically brown or black, with a clear, gelatine like, covering.

The eggs will hatch after a short time, releasing tadpoles. The tadpoles are entirely aquatic, and undergo the most develpoment in the life cycle. They will undergo metamorphisis, in which they will develop legs, and lungs, to become a froglet, resembling an adult but retaining a vestigial tail. Finally the froglet develops into an adult frog. Typically, tadpoles are herbivores, feeding mostly on algae, whereas juvenile and adult frogs are rather voracious carnivores.

Most temperate species of frog reproduce in the period between late autumn to early spring. In the UK most common frog populations produce frogspawn in February although there is wide variation in timing. Water temperatures at this time of year are relatively low and typically between four and 10 degrees celsius. Reproducing in these conditions helps the developing tadpoles because dissolved oxygen concentrations in the water are highest at cold temperatures. More importantly, reproducing early in the season ensures that appropriate food is available to the developing frogs at the right time.

Diet and Predators

All adult frogs are carnivores, and usually eat invertebrates such as: insects, worms and spiders. However, a few of the larger species may eat larger prey, such as small mammals, fish and smaller frogs. Some frogs use their sticky tongues effectively in catching fast-moving prey, while others capture their prey, and force it into their mouth with their hands.

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